Stay duration (means ± SEM, in minutes) in each quadrant of the arena, in the groups that first underwent two-wall shift and then translocation (a) and the groups that first underwent four-wall shift and then removal (b). Environmental setting is depicted at the x-axis and quadrants labeling at the top left. In two-wall shift, rats remained in quadrant 1 (which is equal to the entire arena before expansion). When objects were then translocated, the rats split their time between the starting quadrant (1) and the quadrant that now contained the object array (4). In four-wall shift the rats spent most of the time in quadrant 1, which contained both the start corner and the bottom left object. This pattern was not affected when the objects were then removed, indicating that the rats kept traveling in the same quadrant (see Figure 7).