Joining complexes. (A) Graph representation of two optimized complexes (n = 8 each), joined by reciprocal "inter-modular" connections (stippled arrows). For simplicity, these connections are arranged as n pairs of bi-directional connections, representing a simple topographic mapping. (B) Connection matrix of a network of size n = 16 constructed from two smaller optimized components (n = 8). (C) Information integration as a function of inter-modular coupling strength. Connection strength is given as inter-modular CONij values; all intra-modular connections per element add up to 0.5-CONij. Plot at the bottom shows the size of the complex with highest Φ as a function of inter-modular coupling strength.